COIMBATORE, (India): Twenty-eight elephants have been tested for Covid-19 at a forest reserve in southern India, officials said Wednesday, after the reported death of a rare Asiatic lion from the virus.

In what is believed to be the first known death of an animal in India from the coronavirus, a nine-year-old lioness at a zoo in Chennai in Tamil Nadu state passed away in early June, local media reported.

The feline was among nine lions that had tested positive for the virus, including two who were in critical condition, Chennai’s The New Indian Express newspaper reported last week.

“After lions in Vandalur Zoo tested positive for Covid-19, we were asked to test the camp elephants as a precautionary measure,” a forest ranger at Mudumalai Tiger Reserve in the south of Tamil Nadu told AFP on Wednesday.

Nasal and anal samples were taken from 28 elephants, including two calves, on Tuesday, and sent to the Indian Veterinary Research Institute in the northern state of Uttar Pradesh. The animals’ handlers lifted their trunks to collect a sample from a nostril, according to video shared by the reserve. They also inserted swabs into the elephants’ rectums.